Jessie's father is murdered and Lena, her sister, is collateral damage (it is thought to be an accidental gas leak). Jessie and her mother Anna move near Kylie and her mother Tanya. Anna wants Kylie as a replacement.

When Rumpelstiltskin destroys the Magic Mirror and escapes to the modern world, the four princesses of "Once Upon a Time"-Cinderella, Sleeping Beauty, Snow White, and Rapunzel-are sucked through the portal too. Well-trained and endowed with magical powers, the four women must fight Rumpelstiltskin and his army of thralls before he enslaves everyone on Earth.